A Model of Lysosomal pH Regulation
- Journal of General Physiology 141 (2013): 705-720. With Smita Nayak, Joseph A. Mindell, and Michael Grabe [Abstract and PDF]
Sewall Wright and Gustave Malécot on Isolation by Distance
- Philosophy of Science 76 (2009): 784-796. [Abstract and PDF]
Transposable Elements and an Epigenetic Basis for Punctuated Equilibria
- BioEssays 31 (2009): 715-726. With David Zeh (first author) and Jeanne Zeh. [Abstract and PDF]
Patterns, Models, and Predictions: Robert MacArthur's Approach to Ecology
- Philosophy of Science 74 (2007): 642-653. [Abstract and PDF]