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- Large, enveloped, bullet-shaped, RNA virions (~60 x 180nm) with short glycoprotein spikes.

- Moderately resisitant; survive well in dark places at low temperatures (burrows, caves, soil).

- Includes a wide range of pathogens infecting many species eg: Vesicular Stomatitis Virus (cannot be differentiated clinically from foot-and-mouth disease virus; insect transmission); Ephemeral Fever (infection of tropical cattle; insect transmission); Fish Rhabdoviruses (mortality often high where intensive fish farming practised).