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The principal viral cause of the SMEDI syndrome in the U.K. (stillbirths, mummification and embryonic deaths). Replicates in the epithelium of the alimentary tract with viraemia without causing clinical signs in fattening and breeding stock. Virus recovered from most excretions. Transplacental infection occurs. In early pregnancy, causes resorption and may maifest as infertility; later in pregnancy, SMEDI syndrome. Congenitally infected pigs may remain as excretors; some piglets in litter escape infection.