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Before then in the 1880s, Walter Flemming (1843-1905) developed mitosis, and meiosis was developed by Edouard Van Beneden (1846-1910). In 1905, Nettie Stevens (1861-1912) observed in Tenebrio beetles that all pairs of homologous chromosomes match, except for one pair of X & Y which determines sex.In the same year, 1909, Edmund Beecher Wilson (1859-1939) also established the sex chromosomes. Hugo De Vries (1848-1935) interpreted the works of Gregor Johan Mendel (1822-1884) and the theory of inheritance.